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Raymond A. CloydProfessor 239 W. Waters Hall Manhattan KS 66506 Phone: 785-532-4750 Fax: 785-532-6232 |
Area(s) of Specialization:
Integrated Pest Management, Plant Protection, Horticultural and Ornamental Cropping Systems, Biological Control, Insect-Plant Interactions, Pesticides
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General Statement
Dr. Raymond A. Cloyd has an extension (70%) and research (30%) appointment at Kansas State University (Manhattan, KS). His research and extension program involves pest management/plant protection in greenhouses, nurseries, landscapes, turfgrass, conservatories, interiorscapes, Christmas trees, and vegetables and fruits. Dr. Cloyd is the extension specialist in horticultural entomology for the state of Kansas with a major clientele that includes homeowners, master gardeners, and professional and commercial operators.
Education
B.S. Ornamental Horticulture, California Polytechnic State University -San Luis Obispo, 1990
M.S. Entomology, Purdue University, 1995
Ph.D. Entomology, Purdue University, 1999
Research Program
Dr. Cloyd's research interests are associated with pest management/plant protection, biological control, plant-insect interactions, and non-chemical means of dealing with insect and mite pests. His research program involves pest management/plant protection of insect and mite pests of horticultural crops including thrips, fungus gnats, mealybugs, whiteflies, and spider mites. His research emphasis is on scouting, sanitation, biological control, and appropriate use of pest control materials (insecticides and miticides). His current research projects affiliated with greenhouse pest management are 1) biological control of fungus gnats (Bradysia spp.) using the rove beetle, Dalotia coriaria; 2) effect of systemic insecticides on the citrus mealybug (Planococcus citri); and 3) use of entomopathogenic fungi in rotation programs to mitigate populations of the western flower thrips (Frankliniella occidentalis).
Dr. Cloyd also has existing research projects in turfgrass, nurseries, conservatories, and vegetables.
Publications
A. Scientific (2009 to 2013)
Cloyd, R. A. Winter 2013. Secrets of the "big bugs" special effects in 1950s science fiction movies. American Entomologist 59(4): 203-207.
Hogendorp, B. K., and R. A. Cloyd. 2013. Effect of potassium bicarbonate (MilStop®) and insecticides on the citrus mealybug, Planococcus citri (Risso), and the natural enemies Leptomastix dactylopii (Howard) and Cryptolaemus montrouzieri (Mulsant). HortScience 48(12): 1513-1517.
Echegaray, E. R., and R. A. Cloyd. 2013. Life history characteristics of the rove beetle, Dalotia
coriaria (Coleoptera: Staphylinidae) under laboratory conditions. J. Kansas Entomol. Soc.
86(2): 145-154.
Willmott, A. L., R. A. Cloyd, and K. Y. Zhu. 2013. Efficacy of pesticide mixtures against the
western flower thrips (Thysanoptera: Thripidae) under laboratory and greenhouse conditions.
J. Econ. Entomol. 106(1): 247-256.
Echegaray, E. R., and R. A. Cloyd. 2012. Effects of reduced-risk pesticides and plant growth
regulators on rove beetle (Coleoptera: Staphylinidae) adults. J. Econ. Entomol. 105(6): 2097-
2106.
Hogendorp, B. K., R. A. Cloyd, and J. M. Swiader. 2012. Determination of silicon
concentration in some horticultural plants. HortScience 47(11): 1593-1595.
Johnson, W. A., R. A. Cloyd, J. R. Nechols, K. A. Williams, N. O. Nelson, D. Rotenberg, and
M. M. Kennelly. 2012. Effect of nitrogen source on pac choi (Brassica rapa L.) chemistry
and interactions with the diamondback moth (Plutella xylostella L.). HortScience 47(10):
1457-1465.
Johnson, W. A., J. R. Nechols, R. A. Cloyd, D. Rotenberg, and M. M. Kennelly. 2012. Effect
of light intensity on Brassica rapa chemistry and Plutella xylostella (Lepidoptera:
Plutellidae) life history traits. J. Entomol. Sci. 47(4): 327-349.
Cloyd, R. A., and J. D. Gillespie. 2012. Effect of sugar-based compounds in enhancing the
efficacy of insecticides against the western flower thrips. HortTechnology 22(2): 177-184.
Cloyd, R. A., K. A. Williams, F. J. Byrne, and K. E. Kemp. 2012. Interactions of light
intensity, insecticide concentration, and time on the efficacy of systemic insecticides in
suppressing populations of the sweetpotato whitefly (Hemiptera: Aleyrodidae) and the citrus
mealybug (Hemiptera: Pseudococcidae). J. Econ. Entomol. 105(2): 505-517.
Gontijo, L. M., J. R. Nechols, D. C. Margolies, and R. A. Cloyd. 2012. Plant architecture and
prey distribution influence foraging behavior of the predatory mite Phytoseiulus persimilis
(Acari: Phytoseiidae). Exp. Appl. Acarol. 56: 23-32.
Cloyd, R. A., J. A. Bethke, and R. S. Cowles. 2011. Systemic insecticides and their use in
ornamental plant systems. Floriculture Ornamental Biotech. 5(1): 1-9.
Cloyd, R. A., K. A. Marley, R. A. Larson, A. Dickinson, and B. Arieli. 2011. Repellency of
naturally occurring volatile alcohols to fungus gnat Bradysia sp. nr. coprophila (Diptera:
Sciaridae) adults under laboratory conditions. J. Econ. Entomol. 104(5): 1633-1639.
Fry, J. D., and R. A. Cloyd. 2011. Zoysiagrass genotypes differ in susceptibility to the
bluegrass billbug, Sphenophorus parvulus. HortScience 46(9): 1314-1316.
Cloyd, R. A., and J. A. Bethke. 2011. Impact of neonicotinoid insecticides on natural enemies
in greenhouse and interiorscape environments. Pest Manag. Sci. 67: 3-9.
Hogendorp, B. K., J. M. Swiader, and R. A. Cloyd. 2011. Plant alkaline fusion technique
followed by colorimetric procedure for the detection and quantification of total silicon in
ornamental plants. Comm. Soil Sci. Plant Analysis 42: 75-92.
Cloyd, R. A., K. A. Marley, R. A. Larson, and B. Arieli. 2010. Bounce® fabric softener dryer sheets
repel fungus gnat, Bradysia sp. nr. coprophila (Diptera: Sciaridae), adults. HortScience 45(12):
1830-1833.
Cloyd, R. A. 2010. Pesticide mixtures and rotations: Are these viable resistance mitigatingstrategies? Pest Technol. 4(1): 14-18.
Gontijo, L. M., D. C. Margolies, J. R. Nechols, and R. A. Cloyd. 2010. Plant architecture, prey
distribution and predator release strategy interact to affect foraging efficiency of the
predatory mite Phytoseiulus persimilis (Acari: Phytoseiidae) on cucumber. Biol. Control 53:
136-141.
Cloyd, R. A., C. L. Galle, S. R. Keith, and K. E. Kemp. 2010. Effect of fungicides and
miticides with mitochondria electron transport inhibiting activity on the twospotted spider
mite, Tetranychus urticae (Acari: Tetranychidae). HortScience 45(4): 687-689.
Hogendorp, B. K., R. A. Cloyd, C. Xu, and J. M. Swiader. 2010. Effect of silicon-based
fertilizer applications on nymphal development and adult emergence of the greenhouse
whitefly (Hemiptera: Aleyrodidae) feeding on poinsettia. J. Entomol. Sci. 45(2): 150-169.
Hogendorp, B. K., R. A. Cloyd, and J. M. Swiader. 2009. Effect of silicon-based fertilizer
applications on the reproduction and development of the citrus mealybug (Hemiptera:
Pseudococcidae) feeding on green coleus. J. Econ. Entomol. 102(6): 2198-2208.
Cloyd, R. A. 2009. Western flower thrips (Frankliniella occidentalis) management on ornamental
crops grown in greenhouses: Have we reached an impasse? Pest Technol. 3(1): 1-9.
Hogendorp, B. K., R. A. Cloyd, and J. M. Swiader. 2009. Silicon-based fertilizer applications
have no effect on the reproduction and development of the citrus mealybug, Planococcus
citri Risso (Hemiptera: Pseudococcidae), feeding on fiddleleaf fig, Ficus lyrata (Warb.).
HortScience 44(6): 1616-1621.
Cloyd, R. A., N. R. Timmons, J. M. Goebel, and K. E. Kemp. 2009. Effect of pesticides on adult
rove beetle Atheta coriaria (Coleoptera: Staphylinidae) survival in growing medium. J. Econ.
Entomol. 102(5): 1750-1758.
Cloyd, R. A., K. A. Marley, R. A. Larson, and B. Arieli. 2009. Attractiveness of parboiled rice
hulls to the fungus gnat, Bradysia sp. nr coprophila (Diptera: Sciaridae), adult relative to
standard growing medium components. HortScience 44(5): 1366-1369.
Cloyd, R. A., C. L. Galle, S. R. Keith, N. A. Kalscheur, and K. E. Kemp. 2009. Effect of
commercially available plant-derived essential oil products on arthropod pests. J.
Econ. Entomol. 102(4): 1567-1579.
Cloyd, R. A. 2009. Getting mixed-up: are greenhouse producers adopting appropriate pesticide
mixtures to manage arthropod pests? HortTechnology 19(3): 638-646.
Cloyd, R. A., C. L. Galle, S. R. Keith, and K. E. Kemp. 2009. Evaluation of persistence of
selected miticides against the twospotted spider mite, Tetranychus urticae. HortScience
44(2): 476-480.
Bethke, J. A., and R. A. Cloyd. 2009. Pesticide use in ornamental production: what are the
benefits? Pest Manag. Sci. 65: 345-350.
B. Extension (2009-2013)
Miller, F. L., B. Bauernfiend, R. A. Cloyd, S. Keeley, M. Kennelly, C. Lee, and W. Upham.
September 2013. Turf pest control. Kansas State University Agricultural Experiment Station
and Cooperative Extension Service Category 3B. S-20. Kansas State University, Manhattan,
KS. 64 pgs.
Cloyd, R. A. April 2013. Insect growth regulators for use in greenhouses. Kansas State University
Agricultural Experiment Station and Cooperative Extension Service. MF-3094. Kansas State
University, Manhattan, KS. 2 pgs.
Cloyd, R. A. April 2013. Rose rosette disease. Kansas State University Agricultural Experiment
Station and Cooperative Extension Service. MF-2974. Kansas State University, Manhattan, KS
(revised from March 2011). 2 pgs.
Midwest tree fruit spray guide. 2013. Kansas State University Agricultural Experiment Station and
Cooperative Extension Service S-146. Collaboration with Arkansas, Illinois, Indiana, Iowa,
Kentucky, Missouri, Ohio, and Wisconsin. Entomology: Raymond Cloyd. 68 pgs (revised
annually).
Midwest small fruit and grape spray guide. 2013. Kansas State University Research and Extension.
Collaboration with Arkansas, Illinois, Indiana, Iowa, Kentucky, Missouri, Nebraska, Ohio,
Oklahoma, West Virginia, and Wisconsin. Entomology: Raymond Cloyd. 86 pgs (revised
annually).
Midwest vegetable production guide for commercial growers. 2013. Kansas State University
Research and Extension. Collaboration with Illinois, Indiana, Iowa, Minnesota, Missouri, and
Ohio. Entomology: Raymond Cloyd. 204 pgs (revised annually).
Rebek, E. J., and R. A. Cloyd. 2013. Management of insects and mites in greenhouse floral crops.
Oklahoma Cooperative Extension Service. CR-6718. Division of Agricultural Sciences and
Natural Resources, Oklahoma State University, Stillwater, OK. 8 pgs.
Hale, F. A., and R. A. Cloyd. 2013. Insect and mite management in greenhouses. The University of
Tennessee, Institute of Agriculture. The University of Tennessee Extension PB 1594,
Knoxville, TN. 21 pgs (extensive revision since 2001).
Cloyd, R. A. 2012. Pesticide metabolites. Kansas State University Agricultural Experiment
Station and Cooperative Extension Service. MF-3070. Kansas State University, Manhattan,
KS. 2 pgs.
Cloyd, R. A. 2012. Pesticide mixtures: Understanding their use in horticultural production
systems. Kansas State University Agricultural Experiment Station and Cooperative Extension
Service. MF-3045. Kansas State University, Manhattan, KS. 4 pgs.
Cloyd, R. A. 2012. Management of insects and mites in greenhouse floral crops. Rebek, E. J.
[ed.]. Oklahoma Cooperative Extension Service. CR-6718. Division of Agricultural Sciences
and Natural Resources, Oklahoma State University, Stillwater, OK. 8 pgs.
Cloyd, R., and M. Kennelly. 2011. Interior landscape pest management. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service Category 3C.
S-33. Kansas State University, Manhattan, KS. 35 pgs.
Bauernfeind, B., R. Cloyd, E. DeWolf, W. Fick, D. Jardine, M. Kennelly, D. Peterson, J.
Whitworth, and R. E. Wolf. 2011. Agricultural plant pest control. Kansas State University
Agricultural Experiment Station and Cooperative Extension Service Category 1A. S-19.
Kansas State University, Manhattan, KS. 126 pgs.
Cloyd, R. A. 2011. Mealybug: Management in greenhouses and nurseries. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service. MF-3001.
Kansas State University, Manhattan, KS. 4 pgs.
Cloyd, R. A. 2011. Twospotted spider mite: Management in greenhouses and nurseries.
Kansas State University Agricultural Experiment Station and Cooperative Extension Service.
MF-2997. Kansas State University, Manhattan, KS. 8 pgs.
Cloyd, R. A. 2011. Rose rosette disease. Kansas State University Agricultural Experiment
Station and Cooperative Extension Service. MF-2974. Kansas State University, Manhattan,
KS. 2 pgs.
M. Kushad, E. Wahle, M. Ogutu, R. Weinzierl, M. Babadoost, J. Masiunas, P. Domoto, G.
Nonnecke, D. Lewis, M. Gleason, T. Carey, R. Cloyd, M. Kennelly, J. Strang, T. Jones, R.
Bessin, J. Hartman, M. Warmund, B. Barrett, K. Striegler, M. Kaps, L. Kovacs, M.
Pszolkowski, P. Read, F. Baxendale, J. Chaky, L. Sandell, I. Dami, S. Wright, D. Doohan, C.
Welty, R. Williams, M. Ellis, E. Stafne, P. Mulder, D. Smith, B. Bordelon, R. Foster, J.
Beckerman, S. Weller, J. Jett, J. Baniecki, R. Harbut, B. Smith, P. McManus, D. Mahr, and J.
Colquhoun. 2010. Midwest small fruit and grape spray guide. Bordelon, B., M. Ellis, and C.
Welty [eds.]. Kansas State University, Agricultural Experiment Station and Cooperative
Extension Service. S-145. 74 pgs.
Cloyd, R. A. 2010. Broad mite and cyclamen mite: Management in greenhouses and nurseries.
Kansas State University Agricultural Experiment Station and Cooperative Extension Service.
MF-2938. Kansas State University, Manhattan, KS. 4 pgs.
Cloyd, R. A. 2010. Bulb mites: Management in greenhouses and nurseries. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service. MF-2939.
Kansas State University, Manhattan, KS. 2 pgs.
Cloyd, R. A. 2010. Fungus gnats: Management in greenhouses and nurseries. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service. MF-2937.
Kansas State University, Manhattan, KS. 4 pgs.
Davis, H., J. R. Whitworth, R. A. Cloyd, and C. Otte. 2010. Bed bugs. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service. MF-2926.
Kansas State University, Manhattan, KS. 4 pgs.
Marr, C. W., T. Carey, R. Cloyd, and M. Kennelly. 2010. Kansas Garden Guide. Kansas State
University Agricultural Experiment Station and Cooperative Extension Service. S-51 (Rev.).
Kansas State University, Manhattan, KS. 76 pgs.
Cloyd, R. A. July 2010. Management of insects and mites in greenhouse floral crops. Rebek E.
J., and T. A. Royer [eds.]. Oklahoma Cooperative Extension Service. CR-6718. Division of
Agricultural Sciences and Natural Resources, Oklahoma State University, Stillwater, OK. 8 pgs.
Cloyd, R. A. 2010. Western flower thrips: Management on greenhouse-grown crops. Kansas
State University Agricultural Experiment Station and Cooperative Extension Service. MF-
2922. Kansas State University, Manhattan, KS. 8 pgs.
Cloyd, R. A., and R. S. Cowles. 2009. Resistance management: Resistance, mode of action, and
pesticide rotation. Kansas State University Agricultural Experiment Station and Cooperative
Extension Service. MF-2905. Kansas State University, Manhattan, KS. 8 pgs.
Quinn, J., D. Trinklein, and R. A. Cloyd. 2009. Integrated pest management in greenhouses and
nurseries. University of Missouri Extension. IPM 1026. United States Department of
Agriculture, Cooperative Extension, University of Missouri, Columbia, MO. 8 pgs.
Cloyd, R. A. 2009. Mulches and living organisms. Kansas State University Agricultural
Experiment Station and Cooperative Extension Service. MF-2900. Kansas State University,
Manhattan, KS. 2 pgs.
Cloyd, R. A., J. Quinn, and D. Trinklein. 2009. Integrated pest management in greenhouses and
herbaceous nurseries. Kansas State University Agricultural Experiment Station and
Cooperative Extension Service. MF-2892. Kansas State University, Manhattan, KS. 7 pgs.
Cranshaw, W. S., and R. A. Cloyd. 2009. Fungus gnats as houseplant and indoor pests. Insect
Series Home & Garden, no. 5.5584. Colorado State University, Fort Collins, CO. 3 pgs.