Cloyd, Raymond A.
Professor
239 W. Waters Hall
Manhattan KS 66506
Phone: 785-532-4750
Fax: 785-532-6232
Area(s) of Specialization:
Integrated Pest Management, Ornamental Horticulture, Biological Control
View My Curriculum Vitae
Supplimental to Curriculum Vitae
Education
B.S. Ornamental Horticulture, California Polytechnic State University -San Luis Obispo, 1990M.S. Entomology, Purdue University, 1995
Ph.D. Entomology, Purdue University, 1999
Research
Publications
Cloyd, R. A., K. A. Marley, R. A. Larson, and B. Arieli. 2010. Bounce® fabric softener
dryer sheets repel fungus gnat, Bradysia sp. nr. coprophila (Diptera: Sciaridae),
adults. HortScience 45(12):
1830-1833.
Gontijo, L.M., D.C. Margolies, J.R. Nechols and R.A. Cloyd. 2010. Plant architecture, prey distribution and predator release strategy interact to affect foraging efficiency of the predatory mite Phytoseiulus persimilis (Acari: Phytoseiidae) on cucumber. Biological Control 53:136-141.
Caamano, E.X., R.A. Cloyd, L.F. Solter, and D.J. Fallon. 2008. Quality assessment of two commercially available species of entomopathogenic nematodes: Steinernemafeltiae and Heterorhabditis indica. Horttechnology 18: 84-89.
Cloyd, R.A. and H. Chiasson. 2007. Activity of an essential oil derived from Chenopodium ambrosioides on greenhouse insect pests. Journal of Economic Entomology 100: 459-466.
Cloyd, R.A., A. Dickinson, R.A. Larson and K.A. Marley. 2007. Phototaxis of fungus gnat, Bradysia sp. nr. coprophila (Lintner) (Diptera: Sciaridae), adults to different light intensities. Hortscience 42: 1217-1220
Cloyd, R.A., A. Dickinson, and K.E. Kemp. 2007. Effect of diatomaceous earth and Trichoderma harzianum T-22 (Rifai strain KRL-AG2) on the fungus gnat Bradysia sp. nr. coprophila (Diptera: Sciaridae). Journal of Economic Entomology 100: 1353-1359
Cloyd, R.A., C.L. Galle, and S.R. Keith. 2007. Greenhouse pesticide mixtures for control of silverleaf Whitefly (Homoptera : Aleyrodidae) and twospotted spider mite (Acari : Tetranychidae). Journal of Entomological Science 42: 375-382
Czapar, G.F., M.P. Curry and R.A. Cloyd. 2007. Educational needs and customer service practices of retail stores that sell pesticides in Illinois. HortTechnology 17:115-119
Cloyd, R.A., A. Dickinson, R.A. Larson and K.A. Marley. 2007. Effect of growing media and their constituents on fungus gnat, Bradysia sp. nr. Coprophila (Lintner) adults. Insect Science 14: 467-475.
Goebel, J., K. Williams and R. Cloyd. 2007. Effect of light intensity on the translocation of two systemic insecticides in poinsettia (Euphorbiapulcherrima) and their subsequent efficacy for sweet potato whitefly (Bemisiaargentifold) control. Hortscience 42: 944
Lash, H.E., D.F. Warnock and R.A. Cloyd. 2007. Effect of pesticide mixtures on the survival of the predatory mite Neoseiulus cucumeris (Acarina: Phytoseiidae). Journal of Entomological Science 42: 311-319
Birken, E.M. and R.A. Cloyd. 2007. Food preference of the rove beetle, Atheta coriaria Kraatz (Coleoptera: Staphylinidae) under laboratory conditions. Insect Science 14: 53-56
Cloyd, R. A., and C. A. Casey. 2006. Chapter 10: Insect, mite, and slug management, pp. 115-133. In: Cutting Propagation: A Guide to Propagating and Producing Floriculture Crops. Dole, J. M., and J. L. Gibson [eds.]. Ball Publishing, Batavia, IL. 385 pgs. ISBN no. 1-883052-48-3
Hogendorp, B.K. and R.A. Cloyd. 2006. Insect management in floriculture: how important is sanitation in avoiding insect problems? HortTechnology 16: 633-636
Tiddens, P. T., and R. A. Cloyd. 2006. Susceptibility of three rose genotypes to Japanese beetle (Coleoptera: Scarabaeidae) adult feeding. J. Arboriculture & Urban Forestry 32(3): 108-113.
Cloyd, R. A., C. L. Galle, and S. R. Keith. 2006. Compatibility of three miticides with the predatory mites Neoseiulus californicus McGregor and Phytoseiulus persimilis Athias-Henriot (Acari: Phytoseiidae). HortScience 41(3): 707-710.
Cloyd, R. A. 2006. Compatibility of insecticides with natural enemies to control pests of greenhouses and conservatories. Journal of Entomological Science 41(3): 189-197.
Weinzierl, R., and R. A. Cloyd. 2006. Insect pest management for commercial vegetable crops, pp. 197-234. In: Illinois Agricultural Pest Management Handbook. University of Illinois Extension. College of Agricultural, Consumer, and Environmental Sciences (extensive revision annually since 2001). 356 pgs. ISBN no. 1-883097-55-X
Cloyd, R. A., and A. Dickinson. 2006. Effect of Bacillus thuringiensis subsp. israelensis and neonicotinoid insecticides on the fungus gnat Bradysia sp nr. coprophila (Lintner) (Diptera: Sciaridae). Pest Manag. Sci. 62: 171-177.
Cloyd, R.A. and A. Dickinson. 2006. Effect of insecticides on mealybug destroyer (Coleoptera : Coccinellidae) and parasitoid Leptomastixdactylopii (Hymenoptera : Encyrtidae), natural enemies of citrus mealybug (Homoptera : Pseudococcidae). Journal of Economic Entomology 99: 1596-1604
Hogendorp, B.K., R.A. Cloyd and J.M. Swiader. 2006. Effect of nitrogen fertility on reproduction and development of citrus mealybug, Planococcus citri Risso (Homoptera : Pseudococcidae), feeding on two colors of coleus, Solenostemon scutellarioides L. Codd. Environmental Entomology 35 (2) 201-211
Meers, T.L. and R.A. Cloyd. 2005. Quantitative sampling method for fungus gnat (Diptera : Sciaridae) eggs in soilless growing media. Journal of Economic Entomology 98 (4) 1399-1403
Cloyd, R. A., and P. L. Nixon. 2005. Managing insects and mites on woody ornamental plants, pp. 31-47. In: Commercial Landscape and Turfgrass Pest Management Handbook. University of Illinois Extension. College of Agricultural, Consumer, and Environmental Sciences, University of Illinois, Champaign-Urbana (extensive revision bi-annually since 2000). 113 pgs. ISBN no. 1-883097-50-9
Loughner, R.L., D.R. Warnock and R.A. Cloyd. 2005. Resistance of greenhouse, laboratory, and native populations of western flower thrips to spinosad. Hortscience 40 (1) 146-149
Cloyd, R.A. 2005. Challenges to extension programming. Horttechnology 15 (1) 177-180.
Warnock, D.F. and R.A. Cloyd. 2005. Effect of pesticide mixtures in controlling western flower thrips (Thysanoptera : Thripidae). Journal of Entomological Science 40 (1) 54-66
Blumthal, M.R., R.A. Cloyd, L.A. Spomer, and D.F. Warnock. 2005. Flower color preferences of western flower thrips. Horttechnology 15 (4) 846-853.
Meers, T.L. and R.A. Cloyd. 2005. Egg-laying preference of female fungus gnat Bradysia sp nr. coprophila (Diptera : Sciaridae) on three different soilless substrates. Journal of Economic Entomology 98 (6) 1937-1942
Cloyd, R.A. and A. Dickinson. 2005. Effects of growing media containing diatomaceous earth on the fungus gnat Bradysia sp nr. coprophila (Lintner) (Diptera : sciaridae). Hortscience 40 (6) 1806-1809
Cabrera, A.R., R.A. Cloyd, and E.R. Zaborski. 2005. Development and reproduction of Stratiolaelapsscimitus (Acari : Laelapidae) with fungus gnat larvae (Diptera : Sciaridae), potworms (Oligochaeta : Enchytraeidae) or Sancassania aff. sphaerogaster (Acari : Acaridae) as the sole food source. Experimental and Applied Acarology 36 (1) 71-81
Cabrera, A.R., R.A. Cloyd, and E.R. Zaborski. 2005. Lethal and sub-lethal effects of novaluron (Pedestal((R))) on the soil-dwelling predatory mite, Stratiolaelaps scimitus (Womersley) (Acari : Mesostigmata : Laelapidae), under laboratory conditions. Journal of Entomological Science 40 (1) 47-53
Cloyd, R.A., S.R. Keith and C.L. Galle. 2004. Effect of the insect growth regulator novaluron (Pedestal) on silverleaf whitefly reproduction. HortTechnology 14 (4) 551-554
Zaborski, E.R. and R.A. Cloyd. 2004. Method for quantitative sampling of fungus gnat larvae in soilless growing media. Journal of Economic Entomology 97 (2) 678-686
Kushad, M.M., R. Cloyd and M.B. Babodoost. 2004. Distribution of glucosinolates in ornamental cabbage and kale cultivars. Scientia horticulturae 101 (3) 215-221
Cloyd, R. A., P. L. Nixon, and N. R. Pataky. 2004. IPM for Gardeners: A Guide to Integrated Pest Management. Timber Press, Portland, OR. 204 pgs. ISBN no. 0-88192-647-7
Gill, S. A., and R. A. Cloyd. 2004. Biological control in bedding plants, pp. 409-417. In: BioControl in Protected Culture, K. M. Heinz, R. G. Van Driesche, and M. P. Parrella [eds]. Ball Publishing, Batavia, IL. 552 pgs. ISBN no. 1-883052-39-4
Rothwangl, K.B., R.A. Cloyd and R.N. Wiedenmann. 2004. Effects of insect growth regulators on citrus mealybug parasitoid Leptomastix dactylopii (Hymenoptera : Encyrtidae). Journal of Economic Entomology 97 (4) 1239-1244
Cabrera, A.R., R.A. Cloyd and E.E. Zaborski. 2004. Effects of greenhouse pesticides on the soil-dwelling predatory mite Stratiolaelaps scimitus (Acari : Mesostigmata : Laelapidae) under laboratory conditions. Journal of Economic Entomology 97 (3) 793-799
Cloyd, R.A. and E.R. Zaborski. 2004. Fungus gnats, Bradysia spp. (Diptera : sciaridae), and other arthropods in commercial bagged soilless growing media and rooted plant plugs. Journal of Economic Entomology 97 (2) 503-510
Sadof, C.S., J.J. Neal and R.A. Cloyd. 2003. Effect of variegation on stem exudates of coleus and life history characteristics of citrus mealybug (Hemiptera : Pseudococcidae). Environmental Entomology 32 (3) 463-469
Cloyd, R.A. 2003. Effect of insect growth regulators on citrus mealybug [Planococcus citri (Homoptera : Pseudococcidae)] egg production. Hortscience 38 (7) 1397-1399
Cabrera, A.R., R.A. Cloyd and E.R. Zaborski. 2003. Effect of monitoring technique in determining the presence of fungus gnat, Bradysia spp. (Diptera : Sciaridae), larvae in growing medium. Journal of Agricultural and Urban Entomology 20 (1) 41-47
Cloyd, R.A. and C.S. Sadof. 2003. Seasonal abundance and the use of an action threshold for western flower thrips, in a cut carnation greenhouse. Horttechnology 13(3) 497-500
Cloyd, R.A. and N.L. Cycholl. 2002. Phytotoxicity of selected insecticides on greenhouse-grown herbs. Hortscience 37 (4) 671-672
Cloyd, R.A. and N.L. Cycholl. 2002. Phytotoxicity of selected insecticides on greenhouse-grown herbs. Hortscience 37 (4) 671-672
Cloyd, R.A., D.F. Warnock and K. Holmes. 2001. Technique for collecting thrips for use in insecticide efficacy trials. Hortscience 36 (5) 925-926
Cloyd, R.A. and C.S. Sadof. 2000. Effects of plant architecture on the attack rate of Leptomastix dactylopii (Hymenoptera : Encyrtidae), a parasitoid of the citrus mealybug (Homoptera : Pseudococcidae). Environmental Entomology 29 (3) 535-541